6): (1) sexual rest, (2) early oogenesis, (3) late oogenesis, (4) maturation, (5) spawning and post spawning.
Stage I: sexual rest, the gonad is loose; connective tissue is very abundant, with the appearance of fewer oogonianot exceeding 6 mm in diameter.
In other hand, the peak observed in January (period of sexual rest), and the increase in spring did not match with the gametogenic cycle.
marginatus according to the histological characteristics of the gonads: 0, sexual rest; I, commencement of gametogenesis; II, gametic growth; III, reproduction; and IV, resorption of the gonad.
marginatus in Spain shows a period of sexual rest from August to October, a period of gametogenesis from November to April, and a period of reproduction from May to June.