Sepals triangular-ovate, elongated, abaxially sparsely pubescent, adaxially
densely pubescent, margins denticulate, apex acuminate.
Thoracic axial rings strongly arched, lacking lateral lobes, up to 38% as wide as whole segment in anterior segments and down to 31% in posterior segments; ornamented by small tubercles of varying sizes, gradually fading adaxially
. Articulating half-ring smooth, bounded posteriorly by a deep ring furrow.
Pattern 5--flat or protruded adaxially
and sharply projected abaxially in Anteremanthus, Chronopappus, Heterocoma and Proteopsis with abaxial surface rounded (Fig.
Prickle hairs on the abaxial surface developed; midrib adaxially
grooved, 8-18 mm long, glabrous to lightly puberulent, petiolar glands absent.
Leaves 17 to 20 in number, coriaceous, densely rosulate, suberect, forming a funnelform rosette; sheaths ovate, 12-17 x 9.5-10 cm, inconspicuously lepidote, green toward the apex, strongly coriaceous; blades narrowly subtriangular-lanceolate, not narrowed at the base, 15-24 x 5-6 cm, green, inconspicously and sparsely white-lepidote mainly adaxially
to glabrous, nerved, suberect with recurved apex, apex acuminate.
attached to the rachis, 11 complete and 2 incomplete leaflets can be observed.
Stipes are short, stiff in adult plants, adaxially
canaliculated, moderate to densely scaly with scales similar to those of rhizome (Fig.
Petiole rounded adaxially
; sori concentrated on small apical or lateral lobes 2.
The palpebral furrow is prominent and delimits the eye-ridge posteriorly, dying out shortly adaxially
, also joining a shallow posterior sutural furrow (indicated by row of pits).
M1 separated from M2 by a distinct furrow slightly inclined anteriorly, then strongly curved forward adaxially
. M2 with a strong median tubercle extended slightly backwards above the transverse furrow bounding its posterior margin.
Young stems and abaxial foliar surfaces with well developed, 0.2-1 mm long, wooly, dendritic, rusty brown hairs with well developed axis and moderate number of terete short to moderately long arms (Wurdack 1980, indumentum type 26), persistent; Mature leaves of each pair equal in size, 3.5-5.5 x (8)-11.5-(13) cm, elliptic-lanceolate, cuspidate apically and acute to obtuse basally, 5-nerved, secondary lateral veins 1-2 mm distant chartaceous to coriaceous, entire, with broadly revolute margins, brownish green and glabrous adaxially
, pale rusty-brown abaxially; petioles 18-30 (35) mm long and 1.5-2.5 mm diameter; petiole length to lamina length 1:(2.2)-2.5-(4); lamina length to width (1.7)-2-(3.9):1.