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It was also believed that the amount of chromatin (and its relation to cytoplasm) produced by the presence or absence of the accessory chromosome, would determine maleness or femaleness [see also Jordan (1910)].
(Tettigoniidae); in McClung's words: 'Here [in Mermiria] it is found that one of the tetrads has become united end to end with the accessory chromosome so that the planes of their longitudinal division are coincident'.