Indeed, it is clear that several taxa of Paleogene Proteaceae fossils do exhibit scleromorphy, as has previously been discussed with respect to Banksia, highly reduced leaf types in Lomatia and Lomatia-like species, and Orites species with sclerified hypodermes.
In Australia, with the exception of Banksia and perhaps Lomatia, lineages that now occur in Mediterranean-climate heathlands, and that are sometimes extremely diverse and abundant there, have essentially non-existent fossil records.
Lomatia is a parallel case to that of Orites, but with an even older estimated stem age [49.
A further point is that because the likelihood of fossilisation of any plant organs is extremely low, the relatively high abundance and/or wide spatial and/or temporal distribution of at least Banksia, Orites and Lomatia fossils itself probably reflects that these plants were important components of the vegetation, and had traits that enabled their lineages to become widely established.
Probable crown group Lomatia and Telopea species were also present in the Oligocene, but this is only based on compelling foliar features in common with extant species, and it may not be possible to support this with the evidence of convincing non-homoplasious synapomorphies.
Early Tertiary Lomatia (Proteaceae) macrofossils from Tasmania, Australia.
and fossil Argentine species of Lomatia is quite similar.
assignable to Lomatia occidentalis, including much more complete
Lomatia occidentalis is characterized by the symmetric, deeply
preserved Lomatia characters, allowing the maintenance of the taxonomic
Lomatia patagonica is unquestionably different from the other
Lomatia preferruginea was described by Berry (1938) from the Rio
Roupala patagonica, like Lomatia occidentalis and L.
of Lomatia and Roupala, it differs from all of them.
species and Lomatia patagonica differ in the way that the lamina is