1] forking about at basal 1/4, with about 11 apical cells and 8 subapical cells, claval veins uniting middle of clavus
handlirschi generally has sparse, thick hairs on the embolium, corium, clavus
, membrane and scutellum, whereas the other two species have the short, thick hairs restricted to the embolium and adjacent areas of the corium.
Hemelytron with clavus
exposed, reaching apex of scutellum; lateral margins of corium slightly carinate, parallel, almost extending to posterior margin of connexivum II; membrane lacking veins, reaching to base of seventh abdominal segment (Fig.
Uncus slender and long, sharp apically; tegumen relatively narrow; valva intermediate in size; cucullus arched; harpe slender and curved distally, shorter than uncus, sharp apically and bifurcate basally; sacculus intermediate in size, left processus blunt, right processus pointed; clavus
triangular; saccus relatively broad (Fig.
Forewings dark brown with yellow marginal spots and yellow clavus
with dark brown margin.
1A, 4) and some other Pentatomomorpha and Cimicomorpha resemble those of some more basal clades of the Heteroptera (Enicocephalomorpha, some Dipsocoromorpha, most Gerromorpha) --there is no claval angle, no claval commissure, the outlines of the forewings are smoothly elongate and the forewings overlap freely at the level of about the proximal 1/4-1/2 of the clavus
and corium fused into a coriaceous pad, meeting each other along midline for almost the entire length; hemelytral membrane reduced to small pads, reaching third or fourth abdominal tergum.
Holotype: NMSA 2737, almost complete but somewhat damaged forewing, with fore and hind margins partially, and apical margin entirely, missing, a sub-basal portion of membrane lost, and the fore margin of clavus
overlapping posterior margin of remigium.
1a] deeply convex distad of clavus
, reaching M (Chen et al.
in that they lack rows of punctures on the clavus
and on the R + M vein, referred below as Odoniellina s.
Forewing first vein with 3 setae near base, then 4-6 setae, and then 3 widely spaced setae in the distal half; second vein with 3 setae; clavus
with 3 veinal setae, the distal seta longer than the other two, with a gap between the basal seta and the two distal setae; cilia moderately and irregularly wavy.
with 5 marginal setae and more or less equally long (Fig.
Hemelytra: largely dark golden to reddish-brown; radial vein, somewhat crescentic macula on disk of corium, sub-basal and sub-apical bands on clavus
plus claval suture, all pale to whitish.
schuhi the dark areas of the clavus
and corium are dark brown whereas in T.
debilis Stal --Forewing marked with two brown transverse bands across sub-basally, on nodal line and in clavus
bizonata Matsumura 6.