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Poor hygiene and sanitation led to high rates of diarrhoeal disease and helminthic infections, particularly ankylostomiasis (hookworm).
These contributions included not only his research on filaria, but also discovery of the lung fluke, a parasitic worm in dogs, as well as excursions into many arenas of medicine, including liver abscess, dermatological conditions such as pemphigus contagiosus and tinea imbricata, ankylostomiasis, kalaazar, trypanosomiasis, schistosomiasis, typhoid, diphtheria, smallpox vaccination, leprosy, plague, dengue, beriberi, and sprue.